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By Miyeko Murase

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2-12). In most orchids, the anther opens toward the center of the flower or toward the surface of the column, though the anther of Prescottia may open laterally. In the Spiranthoideae, Neottieae, and Diurideae, the anther is usually firmly attached to a thick filament and it remains in place after the pollinia are removed (Fig. 2-12C, E). In the Orchideae and Diseae, the base of the anther is so completely united with the column that there is no clear boundary between column and anther (Fig. 2-11G).

The division of one population or lineage into two is the usual first step. Geographic isolation is probably the major factor in separating populations, but not necessarily the only type of separation. 2. Speciation. Speciation and separation or isolation may occur simultaneously in special cases, as in allopolyploidy, the doubling of chromosome number in a hybrid. In most cases, though, geographic isolation is thought to precede speciation. Separation does not necessarily lead to speciation. Many isolated populations may merge again with other populations or become extinct.

The arrowhead-shaped leaves of Pachyplectron (B) also look a bit out of place among the orchids, and the deeply lobed leaves of some Acianthus species (F) would seem more at home on a buttercup. ) Page 21 Arrangement In most orchids the leaves are arranged distichously, or in two ranks, with the leaves alternating on opposite sides of the stem (Fig. 2-2H, N). In many cases the stem or pseudobulb bears only a single leaf, but if we check the orientation of scale leaves and sheaths, we find them to be distichous.

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